Triticale exemplifies man-made intergeneric cereal synthesised by crossing wheat Triticum aestivum or Triticum durum with rye Secale cereale, two distinct genera within tribe Triticeae. Primary hybrid from this wide cross is haploid or triploid with unpaired chromosomes leading to sterility due to meiotic irregularities and univalents. To restore fertility, seedlings treated with colchicine alkaloid that inhibits microtubule polymerization preventing spindle formation, inducing chromosome doubling to create autoallohexaploid 6x or octoploid 8x with complete homologous pairing. This amphiploid
Ref:
Oettler Triticale breeding Adv Agron; Linde-Laursen Genomic in situ Triticale; Singh BD Intergeneric hybrid example.