Mature pollen grains contain:
Mature pollen grains at anthesis contain distinctive cellular composition: one large vegetative cell with diffuse chromatin, abundant organelles, mitochondria, dictyosomes, and extensive cytoskeletal machinery responsible for polarized tube growth, and one smaller generative cell with highly condensed chromatin representing germ lineage enclosed within vegetative cytoplasm. This two-celled arrangement ensures coordinated transport of sperm progenitors alongside vegetative cell controlling navigation through style transmitting tissue. Upon germination vegetative nucleus leads tube elongation wh
Ref: McCormick, Plant Cell 1993; Twell et al.: mature pollen contains one vegetative and one generative cell at dispersal.