Early-successional species are typically selected for rapid occupation of recently disturbed, well-lit habitats. They often produce many small, widely dispersed seeds, germinate quickly, grow rapidly, mature early, and complete relatively short life cycles. Shade intolerance is common because their high photosynthetic capacity performs best in open conditions, while dense later canopies suppress them. Slow growth therefore conflicts with the standard pioneer syndrome and is more characteristic of many late-successional species that conserve resources, tolerate shade, and invest in long-lived t
Ref:
Ecology: Concepts and Applications, Molles, 9th Ed., Ch. 20