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In gymnosperms, the microspore mother cell undergoes meiosis to produce haploid pollen grains.
Ref: NCERT Class 11 Biology Chapter 3: Plant Kingdom Gymnosperms - Characteristics and Life Cycle
The innermost layer is the Tapetum. Its primary function is to nourish the developing pollen grains. Tapetal cells typically possess dense cytoplasm and generally become bi-nucleate or polyploid due to endomitosis or free nuclear division without cytokinesis.
Ref: Cross-section of a microsporangium highlighting the Tapetum layer with dense cytoplasm and multiple nuclei.
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