The character regulated by a holandric gene
metabolic engineering and inheritance analysis shows exogenous gene introduction and pathway extension enable trait improvement. Consequently passes from the father through the son to the grandson. emerges as the valid choice since it demonstrates metabolic extension and combinatorial biosynthesis, aligning with established principles in genetics and biotechnology literature.
Ref: Pierce, Genetics: A Conceptual Approach, 6th Edition, Chapter 3, discusses genotype-phenotype correlation and inheritance patterns providing framework for genetic transmission and metabolic engineering.
