Vertical interrupted annulus is typical of
Advanced Polypodiales including Polypodiaceae, Pteridaceae, Aspleniaceae exhibit sporangia with vertical incomplete annulus interrupted by stomium lips and subtended by long slender stalk, enabling classic elastic ejection with coiling and sudden recoil in dry conditions. Primitive ferns like Osmunda have subterminal transverse annuli, Gleicheniaceae oblique, clubmosses lack annulus entirely having simple longitudinal dehiscence, horsetails possess sporangiophores with no annulus but spores with elaters. Vertical interrupted type reflects derived condition optimizing explosive mechanism and sy
Ref: Kramer & Green 1990; Tryon & Lugardon Spores of Pteridophyta; Advanced Ferns