The model where early colonizers have no effect on later species is:
In the tolerance model, early colonists have little direct effect on recruitment of later species. Different species may enter the site, but those able to tolerate progressively lower resources or altered conditions survive and eventually dominate; replacement does not require facilitation by pioneers. Classical succession terminology divides community development into site exposure, arrival, establishment, interaction, environmental reaction, and relative stabilization. Modern ecology treats these as useful descriptions rather than a rigid deterministic program. Local resource conditions, dis
Ref: Ecology: Concepts and Applications, Molles, 9th Ed., Ch. 20