Which model component represents handling time?
In optimal foraging notation, h denotes handling time: the interval required to pursue, subdue, process, and consume a prey item after encounter. Profitability is often expressed as E/h, so the same energetic reward becomes less attractive as handling time increases. S conventionally denotes search time, E the energetic return, and P may represent encounter probability, prey density, or another model-specific parameter. Distinguishing search from handling is important because a predator can change diet breadth when preferred prey become difficult to find even if their handling profitability remains high. The symbol h specifically captures the post-encounter time cost attached to a prey item. A careful interpretation retains the assumptions of the underlying model and avoids extending it beyond available evidence. Within those assumptions, the keyed concept gives the most consistent account of the biological pattern and its expected outcome. This interpretation follows ecological definitions based on effects on fitness, energy flow, behavior, and population performance. It also shows why superficially similar alternatives can represent different mechanisms once the direction of benefit, harm, or resource transfer is considered.
Ref: Ecology: From Individuals to Ecosystems, Begon et al., 5th Ed., Ch. 10