Which model component represents handling time?
In optimal foraging notation, h denotes handling time: the interval required to pursue, subdue, process, and consume a prey item after encounter. Profitability is often expressed as E/h, so the same energetic reward becomes less attractive as handling time increases. S conventionally denotes search time, E the energetic return, and P may represent encounter probability, prey density, or another model-specific parameter. Distinguishing search from handling is important because a predator can change diet breadth when preferred prey become difficult to find even if their handling profitability re
Ref: Ecology: From Individuals to Ecosystems, Begon et al., 5th Ed., Ch. 10