Complementarity raises ecosystem function when species differ in resource use, timing, rooting depth, microhabitat, or facilitative effects. As richness increases, these distinct contributions accumulate, enabling more complete resource capture or mutually beneficial interactions than a single species could achieve. Species richness counts taxa but does not show their abundance, traits, or interaction strengths. Dominant species may control bulk process rates, rare species may provide specialized functions or future insurance, and predators can restructure whole food webs through indirect effe
Ref:
Ecology: Concepts and Applications, Molles, 9th Ed., Ch. 20