In a species-interaction sign matrix, (+,-) denotes exploitation such as predation or parasitism, because one participant gains while the other loses. (-,-) denotes competition, (+,+) denotes mutualism, and (+,0) denotes commensalism. Consequently, the sequence maps to predation, competition, mutualism, and commensalism, which is listed in choice A. The workbook key C reverses the first two relationships and labels (+,0) as amensalism; amensalism is instead (-,0). This is an indisputable key mismatch under standard ecological notation. Sign order can swap the identities of participants, but it cannot turn a positive-negative outcome into competition or a positive-zero outcome into amensalism. This interpretation follows ecological definitions based on effects on fitness, energy flow, behavior, and population performance. It also shows why superficially similar alternatives can represent different mechanisms once the direction of benefit, harm, or resource transfer is considered. The distinction is biologically useful because ecological labels summarize mechanisms that generate testable predictions. Evaluating costs, benefits, timing, and the identities of interacting organisms prevents confusion between terms that may look similar in a short description.
Ref:
Ecology: From Individuals to Ecosystems, Begon et al., 5th Ed., Ch. 8-13