Hydrarch, or hydrosere, succession begins in a freshwater habitat such as a pond and, under suitable regional climate, proceeds toward a terrestrial community. Phytoplankton and submerged plants trap sediment and add organic matter. As the basin becomes shallower, floating plants, reeds, and sedges colonize; marsh and swamp stages may follow. Continued sedimentation, peat formation, transpiration, and soil development reduce open water and permit shrubs and trees. In a humid climate capable of supporting woodland, the conventional textbook endpoint is forest. A pond is the starting habitat, while marsh and swamp are intermediate stages rather than the final climatic community. This progression is neither inevitable nor rapid: floods, fire, grazing, human management, nutrient supply, and basin hydrology can arrest or reverse it, and a dry regional climate would produce a different endpoint. Modern succession theory also avoids treating every sequence as a fixed march to one climax. Nonetheless, the keyed forest stage follows the classic hydrosere mechanism of infilling, organic accumulation, and increasing terrestrialization.
Ref:
Ecology: Concepts and Applications, Molles, 9th Ed., Ch. 20