An energy pyramid displays the rate at which energy is fixed and transferred from producers through successive consumer levels. Values should be expressed per unit area and time, allowing direct comparison of trophic production. Width decreases upward because incomplete consumption, egestion, and respiration prevent all lower-level production from becoming higher-level biomass. It therefore differs from biomass and number pyramids, which describe standing stocks and can be inverted. The energy pyramid integrates turnover and exposes the thermodynamic structure of the ecosystem. The ten-percent
Ref:
Fundamentals of Ecology, Odum & Barrett, 5th Ed., Ch. 3