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#Bruce Wallace

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Dominance hypothesis of heterosis was proposed by:

Dominance hypothesis advanced by Davenport in 1908 and expanded by Bruce, Keeble and Pellew and Jones in 1917 postulates that heterosis results from complementation of deleterious recessive alleles carried in different homozygous lines. Inbred parents accumulate different sets of mildly deleterious recessives at many loci encoding enzymes of basic metabolism, photosynthetic electron transport and stress defense. Hybrid receives functional dominant wild-type alleles from each parent at complementary loci, restoring pathway efficiency and increasing fitness. Data from maize whole-genome sequenci

Ref: Davenport CB 1908 Degeneration studies; Jones DF 1917 Dominance of linked factors; Charlesworth Review Genetics dominance heterosis.