An energy pyramid is never inverted under consistent boundaries because each trophic level can convert only a fraction of the preceding level’s production into its own production. This makes its shape more robust than a biomass pyramid, which can change with season, lifespan, and turnover. The deeper mechanism is respiratory heat loss plus incomplete consumption and assimilation; “never inverted” describes the outcome rather than the cause. Rate-based energy measurement therefore avoids misleading snapshots of rapidly renewed producers such as phytoplankton. A snapshot may differ seasonally, e
Ref:
Fundamentals of Ecology, Odum & Barrett, 5th Ed., Ch. 3