Species richness often rises during succession as soil development, structural complexity, and habitat heterogeneity create additional niches and more species reach the site. The trend is a broad expectation rather than an inviolable rule, because strong dominance or late disturbance can produce plateaus or declines. Successional trajectories emerge from dispersal, establishment, species interactions, and organism-driven environmental change. Priority effects can make arrival order important, while retained soil and propagules strongly accelerate recovery after disturbance. Facilitation, toler
Ref:
Ecology: Concepts and Applications, Molles, 9th Ed., Ch. 20